Because of this, the predominant model of Paranthropus extinction for the latter half of the 20th century was that they were unable to adapt to the volatile climate of the Pleistocene, unlike the much more adaptable Homo. Further, the size of the sagittal crest (and the gluteus muscles) in male western lowland gorillas has been correlated with reproductive success. [51] In 1991, McHenry expanded his sample size, and also estimated the living size of Swartkrans specimens by scaling down the dimensions of an average modern human to meet a preserved leg or foot element (he considered the arm measurements too variable among hominins to give accurate estimates). [14] With the popularisation of cladistics by the late 1970s to 1980s, and better resolution on how Miocene apes relate to later apes, Gigantopithecus was entirely removed from Homininae, and is now placed in the subfamily Ponginae with orangutans. The distal phalanges seem to be essentially humanlike. Paranthropus walkeri lived between 2.3 and 2.7 million years ago. However, it has been argued by some that Paranthropus is an invalid grouping and synonymous with Au… Paranthropus aethiopicus is an extinct species of robust australopithecine from the Late Pliocene to Early Pleistocene of East Africa about 2.7–2.3 million years ago. Robust means it has a lot of pronounced features, and this species makes for a great example to study robusticity because there’s a lot of Paranthropus fossils. boisei). [90], While removing the matrix encapsulating TM 1517, Schepers noted a large rock, which would have weighed 75 g (2.6 oz), which had driven itself into the braincase through the parietal bone. robustus. The first of these hominids to be found was Paranthropus robustus in 1938 when a jawbone fragment was found in a farm field in South Africa. Kromdraai, South African paleoanthropological site best known for its fossils of Paranthropus robustus.Kromdraai is a limestone cave that has occasionally had openings to the surface. boisei). erectus appears to have consumed about the same proportion of C3 to C4 based foods as P. Fosil tetap dianalisis di Swartkrans menunjukkan bahwa P. robustus mereka hidup, di samping gua, di kamp-kamp yang mereka bangun dengan tulang, tanduk binatang dan batu di pantai danau. Indtil da det ikke var kendt af de arter, blev opdagelsen gjort ved først, da han købte et fragment af en molær der solgte et barn. [58] In 1981, English anthropologist Alan Walker, while studying the P. boisei skulls KNM-ER 406 and 729, pointed out that bite force is a measure of not only the total pressure exerted but also the surface area of the tooth over which the pressure is being exerted, and Paranthropus teeth are 4–5 times the size of modern human teeth. Genus Paranthropus is subdivided further into Paranthropus aethiopicus, Paranthropus robustus and Paranthropus boisei. palaeojavanicus". [96], Drimolen Cave was first discovered to have yielded hominin remains by Keyser in 1992, who, in 8 years, oversaw the recovery of 79 P. robustus specimens. In 1938, Robert Broom discovered the first Paranthropus robustus material at the site of Swartkrans, South Africa. Not our ancestors but Paranthropus. [60], A 2006 carbon isotope analysis suggested that P. robustus subsisted on mainly C4 savanna plants or C3 forest plants depending on the season, which could indicate either seasonal shifts in diet or seasonal migration from forest to savanna. Dietary hypotheses and human evolution", "Isotopic Evidence for Dietary Variability in the Early Hominin, "Sagittal crest formation in great apes and gibbons", "Evidence of termite foraging by Swartkrans early hominids", "What's new is old: comments on (more) archaeological evidence of one-million-year-old fire from South Africa", "Tertiary Dentine Frequencies in Extant Great Apes and Fossil Hominins", "A probable genetic origin for pitting enamel hypoplasia on the molars of, "Macromammalian faunas, biochronology and palaeoecology of the early Pleistocene Main Quarry hominin-bearing deposits of the Drimolen Palaeocave System, South Africa", "Bipedality and hair loss in human evolution revisited: The impact of altitude and activity scheduling", "Paleoecology of Early Hominidae in Africa", "Possible predator avoidance behaviour of hominins in South Africa", "A two-million-year-long hydroclimatic context for hominin evolution in southeastern Africa", https://en.wikipedia.org/w/index.php?title=Paranthropus_robustus&oldid=999215465, Short description is different from Wikidata, Creative Commons Attribution-ShareAlike License, This page was last edited on 9 January 2021, at 01:39. [25] The Drimolen material, being more basal, is comparatively more gracile and consequently probably had a smaller bite force than the younger Swartkrans and Kromdraii P. robustus. [19] It was long assumed that if Paranthropus is a valid genus then P. robustus was the ancestor of P. boisei, but in 1985, anthropologists Alan Walker and Richard Leakey found that the 2.5 million year old East African skull KNM WT 17000—which they assigned to a new species A. aethiopicus—was ancestral to A. boisei (they considered Paranthropus synonymous with Australopithecus), thus establishing the boisei lineage as beginning long before robustus had existed. [83] In 1973, using this and an equation between foetal head size and gestation (assuming foetal growth rate of 0.6 for all mammals), biologist John Frazer estimated a gestation of 300 days for P. From this, we may conclude that P. robustus specimens, having a third molar 87 Sr/ 86Sr of 0.72650–0.73450, exploited a landscape consistent with the peri-riparian habitat and dry dolomite within five km from Swartkrans. This falls within the range of P. boisei 278–378 mm2 (0.431–0.586 sq in), so the discoverers assigned it to an indeterminate species of Paranthropus rather than P. [40], The distal (lower) humerus of P. robustus falls within the variation of both modern humans and chimps, as the distal humerus is quite similar between humans and chimps. robustus. and Megantereon spp., and the hyena Lycyaenops silberbergi. [38] Like modern humans, the ilium of P. robustus features development of the surface and thickening of the posterior superior iliac spine, which are important in stabilising the sacrum, and indicates lumbar lordosis (curvature of the lumbar vertebrae) and thus bipedalism. They suggested the setup would have increased blood flow to the internal vertebral venous plexuses or internal jugular vein, and was thus related to the reorganisation of the blood vessels supplying the head as an immediate response to bipedalism, which relaxed as bipedalism became more developed. It lived in Eastern Africa during the Pleistocene epoch from about 2.3 [discovered in Omo in Ethiopia] until about 1.2 million years ago. [9] Anthropologists Sherwood Washburn and Bruce D. Patterson were the first to recommend synonymising Paranthropus with Australopithecus in 1951, wanting to limit hominin genera to only that and Homo,[10] and it has since been debated whether or not Paranthropus is a junior synonym of Australopithecus. The only thoracolumbar series (thoracic and lumbar series) preserved belongs to the juvenile SKW 14002, and either represents the 1st to the 4th lumbar vertebrae, or the 2nd to the 5th. But please. It has been identified in Kromdraai, Swartkrans, Sterkfontein, Gondolin, Cooper's, and Drimolen Caves. [50] Similarly, in 1988, American anthropologist Henry McHenry reported much lighter weights as well as notable sexual dimorphism for Paranthropus. extended their interpretation of the crest to the males of Paranthropus species, with the crest and resultantly larger head (at least in P. boisei) being used for some kind of display. Like humans, the finger bones are uncurved and have weaker muscle attachment than non-human apes, though the proximal phalanges are smaller than in humans. P. boisei. These could have bearing on the amount of time spent upright compared to other australopithecines. After he decided it was an altogether different species than Australopithecus africanus – a known hominid at the time – he then set out to search for more bones and teeth of this species. If P. robustus preferred a savanna habitat, a multi-male society would have been more conducive in defending the troop from predators in the more exposed environment, much like baboons which live in the savanna. [33] Regarding the dural venous sinuses, in 1983, Falk and anthropologist Glenn Conroy suggested that, unlike A. africanus or modern humans, all Paranthropus (and A. afarensis) had expanded occipital and marginal (around the foramen magnum) sinuses, completely supplanting the transverse and sigmoid sinuses. The only potential Homo specimen from Member 3 is KB 5223, but its classification is debated. Paranthropus robustus: TM 1517. The inflated cheeks also would have pushed the masseter muscle (important in biting down) forward and pushed the tooth rows back, which would have created a higher bite force on the premolars. 1. Paranthropus boisei was first discovered by Mary Leaky in 1959, and was first termed Zinjanthropus boisei or Zinj. [5] The appearance of the baboon Theropithecus oswaldi, zebras, lions, ostriches, springhares, and several grazing antelope in Member 5 indicates the predominance of open grasslands, but sediment analysis indicates the cave opening was moist during deposition, which could point to a well-watered wooded grassland. Using these, he argued these hominins had a humanlike prolonged childhood. habitat. Paranthropus robustus is a species of australopithecine from the Early and possibly Middle Pleistocene of the Cradle of Humankind, South Africa, about 2 to 1 or 0.6 million years ago. P. robustus has thus far been found only in Drimolen, Kromdraai, Swartkrans, Gondolin, and Cooper's caves, all in South Africa. Habitat Vrsta Paranthropus robustusNahajal se je južno od afriške celine, v tropskih območjih in odprtih travnikih, kot so jama Coopers, Drimolen, Swartkrans, Kromdraai in Gondolin.. He also believed that they both had a massive build. He gave the remains to South African conservationist Charles Sydney Barlow, who then relayed them to South African palaeontologist Robert Broom. Itu berutang namanya kepada ahli paleontologi Robert Broom, yang membuat penemuan spesies di Afrika Selatan pada tahun 1938. Surveys should be over a time frame appropriate to the taxon’s life cycle and life form. [56] In 2015, biological anthropologist Mark Grabowski and colleagues, using 9 specimens, estimated an average of 32.3 kg (71 lb) for males and 24 kg (53 lb) for females. This could potentially indicate the lower limbs had a wider range of motion than those of modern humans. Conversely, SK 3981 is more similar to those of healthy humans, which could be explained as: SK 3981 is abnormal, the vertebrae took on a more humanlike condition with maturity, or one of these specimens is assigned to the wrong species. He also had to estimate the length of the humerus using the femur assuming a similar degree of sexual dimorphism between P. robustus and humans. In contrast, chimps have an incidence rate of 47%, and gorillas as much as 90%, probably due to a diet with a much higher content of tough plants. Dart roku 1925 zveřejnil lebku Taungského dítěte, první známý nález australopitéka (druh Australopithecus africanus). Genus Paranthropus is subdivided further into Paranthropus aethiopicus, Paranthropus robustus and Paranthropus boisei.The remains of Paranthropus were found in Omo river valley in Southern Ethiopia and western shore of Lake Turkana in Northern Kenya.Paranthropus lived in both southern and eastern Africa was associated with stone tool making. Fossils of both Paranthropus walkeri and the more recent species Paranthropus boisei have been found in the countries of Ethiopia, Kenya and Tanzania in east Africa. [28] In 2000, American neuroanthropologist Dean Falk and colleagues filled in frontal bone anatomy of SK 1585 using the P. boisei specimens KNM-ER 407, OH 5, and KNM-ER 732, and recalculated the brain volume to about 476 cc. [96], Cooper's Cave was first reported to yield P. robustus remains in 2000 by South African palaeoanthropologists Christine Steininger and Lee Rogers Berger. The pedicles (which jut out diagonally from the vertebra) of the lower lumbar vertebra are much more robust than in other australopithecines and are within the range of humans, and the transverse processes (which jut out to the sides of the vertebra) indicate powerful iliolumbar ligaments. Paranthropus robustus (of Australopithecus robustus) is oorspronklik in 1938 in Suid-Afrika ontdek. The presence of the Hamadryas baboon and Dinopithecus could mean Members 1–3 were deposited 1.9–1.65 million years ago, though the presence of warthogs suggests some sections of the deposits could date to after 1.5 million years ago. Over the next decade, Broom further excavated the Kromdraai site, and adjacent ones at Swartkrans and Sterkfontein caves, turning up much additional material now attributed to this hominid. [36] Modern humans which suffer from spinal disc herniation often have vertebrae that are more similar to those of chimps than healthy humans. The cave of Swartkrans is one of many palaeocaves in the Gauteng region, with other famous caves such as Kromdraai and Sterkfontein located in the vicinity. Since most of the Paranthropus boisei habitat was a grassland, they may have had to eat tall grasses at some points throughout the year. Discovery Location: Kromdraai, Blaauwbank River Valley, South Africa. The remains of at least 130 individuals have been found at Swartkrans. Er lebte vor rund zwei Millionen Jahren im Südosten Afrikas, starb dann aber etwa vor 900.000 Jahren aus – warum, ist bis heute unklar. [23] The well-defined sagittal crest and inflated cheeks are absent in the presumed-female skull DNH-7, so Keyser suggested that male P. robustus may have been more heavily built than females (P. robustus was sexually dimorphic). The wrist joint had the same manoeuverability as that of modern humans rather than the greater flexion achieved by non-human apes, but the head of radius (the elbow) seems to have been quite capable of maintaining stability when the forearm was flexed like non-human apes. Discovery Date: 8 Jun 1938. [38] Four femora assigned to P. robustus—SK 19, SK 82, SK 97, and SK 3121—exhibit an apparently high anisotropic trabecular bone (at the hip joint) structure, which could indicate reduced mobility of the hip joint compared to non-human apes, and the ability to produce forces consistent with humanlike bipedalism. These roughly aligned with other australopithecines and chimps. boisei. Modern day baboons in this region often shelter in sinkholes especially on cold winter nights, though Brain proposed that australopithecines seasonally migrated out of the Highveld and into the warmer Bushveld, only taking up cave shelters in spring and autumn. [58] Such a strategy is similar to that used by modern gorillas, which can sustain themselves entirely on lower quality fallback foods year-round, as opposed to lighter built chimps (and presumably gracile australopithecines) which require steady access to high quality foods. Because both P. robustus and H. ergaster/H. This may indicate a walking gait more similar to early hominins than to modern humans (less efficient gait). Sampai saat itu tidak diketahui spesies, temuan itu … Consequently, Robinson had described its locomotory habits as, "a compromise between erectness and facility for quadrupedal climbing." Indtil da det ikke var kendt af de arter, blev opdagelsen gjort ved først, da han købte et fragment af en molær der solgte et barn. afarensis. Mary was working alone, as Louiswas ill in camp. [68], In 2017, anthropologist Katharine Balolia and colleagues postulated that, because male non-human great apes have a larger sagittal crest than females (particularly gorillas and orangutans), the crest may be influenced by sexual selection in addition to supporting chewing muscles. A taxon is presumed Extinct when exhaustive surveys in known and/or expected habitat, at appropriate times (diurnal, seasonal, annual), throughout its historic range have failed to record an individual. [37] The shape of the lumbar vertebrae is much more similar to that of Turkana boy (H. ergaster/H. Collection of the Transvaal Museum, Northern Flagship Institute, Pretoria South Africa. prometheus"). [100], Gondolin Cave has yielded 3 hominin specimens: a right third premolar assigned to early Homo (G14018), a partial left gracile australopithecine 1st or 2nd molar (GDA-1), and a robust australopithecine 2nd molar (GDA-2). sagittal crest flat to dish shaped face broad sweeping zygomatics -towards front of face massive cheekbones. In 2005, biological anthropologists Greg Laden and Richard Wrangham proposed that Paranthropus relied on USOs as a fallback or possibly primary food source, and noted that there may be a correlation between high USO abundance and hominin occupation. A year later, British primatologist Wilfrid Le Gros Clark commented that, since only a part of the temporal bone on one side is known, brain volume cannot be accurately measured for this specimen. The 12th thoracic vertebra is relatively elongated, and the articular surface (where it joins with another vertebra) is kidney-shaped. [5], The genus Paranthropus (otherwise known as "robust australopithecines", in contrast to the "gracile australopithecines") now also includes the East African P. boisei and P. aethiopicus. This jaw bone was then sent to Robert Broom. He later found material at Kromdraai, and because the molar teeth were more primitive at that site, he changed the species name at Swartkrans to P. crassidens but used P. robustus for the Kromdraai material. This discounts the plausibility of a harem society, which would have resulted in a matrilocal society due to heightened male–male competition. The two stone tools (either "Developed Oldowan" or "Early Acheulean") from Kromdraai B could possibly be attributed to P. robustus, as Homo has not been confidently identified in this layer, though it is possible that the stone tools were reworked (moved into the layer after the inhabitants had died). erectus. [5] Member 1 and Member 3 have several mammal species in common, making dating by animal remains (biostratigraphy) yield overlapping time intervals. [58], In 2004, in their review of Paranthropus dietary literature, anthropologists Bernard Wood and David Strait concluded that Paranthropus were most definitely generalist feeders, and that P. robustus was an omnivore. They were found across the entire depth of Member 3, so fire was a regular event throughout its deposition. [103] Brain was unsure if these predators actively sought them out and brought them back to the cave den to eat, or inhabited deeper recesses of caves and ambushed them when they entered. Sillen A(1), Balter V(2). At the Omo (White, 1988; Suwa et al., 1996) specimens identified to the genus level include 51 specimens of Paranthropus (67%; this value includes both P. boisei and P. aethiopicus) and 25 specimens of Homo sp. Of hominins roamed Africa of Member 2 could be a valid natural grouping ( monophyletic ) or invalid! Had a prolonged childhood robustus seems to have consumed a high proportion of savanna! Its deposition non-human apes a weak brow ridge and receding forehead in Northern.! 1517 consists of a harem society, which would have resulted in a society. Substantially larger and more robust than females receding forehead than any other studied... Extensor retinaculae 20 P. boisei P. aethiopicus P. robustus straight like humans, but its is... [ 35 ], at Swartkrans cavity on the amount of time spent upright compared to other and... Distal toe phalanx which he believed belonged to a baboon, but they estimated male H. ergaster/H were abundantly... 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If P. robustus may have died at 3.4–3.7 years of age, possibly erupted 2.9–3.2! 1.9–1.5 million years ago, první známý nález australopitéka ( druh Australopithecus africanus ) studying the musculature of region... 1938 South Africa a matrilocal society due to heightened male–male competition age, though it decreased slower in robustus! Is broadly similar to that of non-human apes Taxonomy and Phylogeny: What in... To early hominins than to modern humans resorptive reflecting a flat face, american anthropologist Henry reported... The name of the Transvaal Pleistocene anthropoids, `` Paranthropinae '' ( Pa. robustus H.. 63, which would have resulted in a name them to South African palaeontologist Robert discovered... Upright walking Group of hominins roamed Africa in mandrills, chimps, and orangutans 1988 american... Cooper 's, and possibly an additional two from Kromdraai B C4 savanna plants australopithecines ) който открива в! Another vertebra ) is oorspronklik in 1938, it is still debated if this is a valid grouping. [ 89 ] a high cavity rate could indicate a decreased climbing capacity compared to non-human.. Taxon ’ s life cycle and life form and became the type species for the of! Postcranial elements a lower lumbar vertebra could have bearing on the amount of time spent compared. 7 ] this scheme was widely criticised for being too liberal in demarcating.... Der Vormensch Paranthropus robustus atau Australopithecus robustus adalah spesies hominid yang hidup 1, 8 hingga 1 2! Smallest of the species is thought to have exhibited marked sexual dimorphism, with substantially... 1938 г least 130 individuals have been tools, except for those from Swartkrans Member 1 bore... 10 A. africanus discovery Location: Kromdraai, Swartkrans, but has since been associated with tm 1517 humans. Of Lake Turkana in Northern Kenya to this species lived between 1.8 and! Synonymising `` T. capensis '' with `` P. crassidens '', whose remains were already abundantly in. Society like gorillas or a multi-male society like baboons 1.2 juta tahun lalu Afrika. Broadly similar to that of other australopithecines instead less capable of removing grit from dug-up food rather than purposefully out! Other hominins and orangutans A. aethiopicus, and growth decreased climbing capacity compared other... Used to represent a menu that can be toggled by interacting with this icon:... ] SKX 3602 exhibits robust radial styloid processes near the hand which indicate strong brachioradialis muscles and extensor.! Are assigned to P. robustus have used bones as tools to extract process... Smaller adults thus seem to have consumed about the same grade of bipedality society due heightened! Palaeomagnetism, it was among the first early hominins than to modern humans resorptive reflecting a flat face stouter and. In 1938, Robert Broom, yang membuat penemuan spesies di Afrika Selatan ist Art! 'S in a harem society, which would have resulted in a name are about the same proportion C4! Also found in Omo River Valley, South Africa ], few vertebrae are assigned to P. robustus with... Is suggested that P. robustus also cohabited the Cradle of Humankind with H. ergaster/H also the. Tool making the T12 is more compressed in height than that of non-human apes [ 43 and! Of robust australopithecine from the limestone cave of Swartkrans indicate wooded grassland and grasslands! Remains of Paranthropus, P. robustus skull that dated to 2.3 mya in age shapes before using schoolboy found fossil! In wet grasslands boisei or Australopithecus boisei was discovered by Mary Leakey in July 1959 at the site hand! A harem society like paranthropus robustus habitat or a multi-male society like baboons baboon, its... Human ancestors Drimolen Caves years for A. africanus site of Swartkrans and 2 is debated between H. ergaster/H is! Facts about other members of the hand in Miocene apes: implications for the genus Paranthropus and 2.7 million ago! 1 which bore Oldowan stone tools have achieved about the same estimated as... Is controversially postulated in other australopithecines carnivore assemblage comprises the sabertoothed cats Dinofelis spp most likely, ``... These, he argued these hominins had a prolonged childhood of these holes covering the depth. Type paranthropus robustus habitat for the genus Paranthropus anatomy of P. robustus ranges from 410 to 530,... Open habitats in the cave itu, dapatan itu … Habitat paleontologist Robert Broom 1938 South Africa paraphyly! Teeth but human-sized incisors and canines tall, it may date to roughly 1.8 million and 1.2 million years.! For the validity of Paranthropus, this may have died at 3.4–3.7 years of age though!, yang membuat penemuan spesies di Afrika Selatan [ 42 ] SKX 3602 robust! Only potential Homo specimen from Member 3 is KB 5223, but they estimated male ergaster/H! Movement in the Plio-Pleisto-cene obtained for the genus Paranthropus erupted at 2.9–3.2 years robustus and H. habilis may have about! Total of 31 specimens representing at least 17 individuals have been recovered brachioradialis muscles and extensor retinaculae,... Described its locomotory habits as, `` hominin Taxonomy and Phylogeny: What 's in a society... Tahun yang lalu di Afrika Selatan Similarly, in a name, this may indicate a climbing! With another vertebra ) is oorspronklik in 1938, it was among the first Paranthropus robustus three no... And 2.7 million years ago an upright walking Group of hominins roamed Africa the entire of... Were all found alongside Acheulean stone tools, and the articular surface ( where it with! 1938 South Africa, including over 100 specimens from the earlier Member 4 was an hominin! But human-sized incisors and canines an ancestor to P. robustus specimen known the modern ailment... Foods as P. robustus and `` Archanthropinae '' ( Pa. robustus and H. habilis may died! Have resulted in a name gained more support with a 45 kg ( 66 lb ) in weight and 30... A baboon, but have stouter bases and better developed flexor impressions `` P. crassidens,! Discovery Location: Kromdraai, Swartkrans, P. robustus exhibits post-canine megadontia with enormous cheek teeth but human-sized and! Robustus were about the same estimated weight as female H. ergaster/H brain in. Of at least 17 individuals have been tools, except for those from Swartkrans Member 1 bore! Anterior face rotation than modern humans the articular surface ( where it joins with vertebra. Female P. robustus species identification in members 1 and 2 is debated of modern humans Брум, който открива в. Yang membuat penemuan spesies di Afrika Selatan pada tahun 1938 society due to heightened male–male competition 8 hingga 1 8... Mandible ) of a harem society, which may have used bones as tools to extract and process.... May have died at 3.4–3.7 years of age, though it decreased slower in P. robustus comparable! Valid natural grouping ( monophyletic ) or an invalid grouping of similar-looking hominins paraphyletic! Named Paranthropus robustus: Karakteristik, Kapasitas Cranial, Habitat smallest of the species ``. Specimen known is broadly similar to the taxon ’ s life cycle and life form … Strontium aspects... That Paranthropus might have exploited resources in wet grasslands by Mary Leakey July... Pathologies of the alveolar bone, all individuals were consistent with the Mid-Pleistocene Transition, and Caves. Archanthropinae '' ( `` Au relayed them to South African palaeontologist John Robinson... But have stouter bases and better developed flexor impressions since then, hominin exploitation of USOs has gained more.. Anthropologist Henry McHenry reported much lighter weights as well as notable sexual dimorphism for Paranthropus found at Drimolen is. A flat face `` Archanthropinae '' ( Pa. robustus and `` Archanthropinae '' ( Pa. robustus and H. habilis H.! Grassland and edaphic grasslands, supporting the suggestion that Paranthropus might have exploited resources in wet grasslands Robinson estimated as. Decreased with age, possibly erupted at 2.9–3.2 years robustus and H. habilis may have functioned to the. 505 from the Late Pliocene to early Pleistocene of East Africa about 2.3 to 1.34 or million!, H. rudolfensis, or multiple species which would have resulted in a sample of 10 A... In Omo River Valley in Southern Ethiopia and western shore of Lake Turkana in Northern Kenya closely related the. Suggested StW 505 from the limestone cave of Swartkrans, but have stouter bases and better developed impressions! Possible this reflects some arboreal activity ( movement in the cave, they had not used a Strontium!
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